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Pipturus ruber

A.Heller (1897)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Rosids Order: Rosales Family: Urticaceae Genus: Pipturus

māmake [mamake], māmaki [mamaki], waimea

Description

Key Characters:

Growth Form: Shrubs 1.5–2 m tall.

Stems: Branches with reddish brown bark, hispid, the hairs ca. 1 mm long, also with red–tipped glands.

Roots:

Leaves: Leaves simple. Alternate, widely spaced. Blades ovate to elliptic, 7.5–18(–25) cm long, (3.5–)4.5–7 cm wide, leaves of young growth up to 31 cm long and 20 cm wide. Apex acuminate. Base rounded to broadly cuneate. Upper surfaces pale green, smooth, veins slightly impressed, intercostal areas flat or slightly raised, sparsely pubescent, glabrate with age; lower surfaces gray to olive gray; blades coriaceous; areoles thinly woolly pubescent. Margins coarsely crenate, apex acuminate. Palmately 3-veined or 5-veined, pinnately veined toward apex, veins bright red, raised, moderately to densely hispid along veins, the hairs spreading, also with numerous red–tipped glands. Petioles (1–)2–7.5 cm long. Stipules present 10–12 mm long, 5–6 mm wide, the veins hispid, persistent.

Flowers: Flowers in clusters 8–9(–12) mm in diameter, staminate flowers few, mixed with pistillate ones. Plants unisexual, apparently gynodioecious (no staminate plants observed). Calyx of staminate flowers herbaceous, 4-lobed, 2–2.2 mm long, long–hirsute; pistillate Calyx red, 1.2–1.3 mm long, enlarging to 1.8–2.2 mm long in fruit, densely long–hirsute. Corolla (petals) absent. Filaments incurved in bud, elastically reflexed when pollen is shed; anthers dithecal, opening by longitudinal slits. Ovary superior (pistillate flowers), psuedomonomerous, 1-celled; ovule 1, basal; style 1. Staminate flower ovary vestigial and sterile.

Fruit: Achenes pale yellow or grayish brown; conical–ovoid; 1.1–1.5 mm long. Seeds 1 per achene.

Ploidy:

Habitat: Mesic valleys; diverse mesic forest; and wet forest.

Elevation Range: 200–1,370 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Kaua'i Endemic

Dispersal Agents


Pollinators

Notes

  • 200-1370 m
  • Description digitized by Olivia Kraus
  • Description digitized from the Manual of the Flowering Plants of Hawaii

Bibliography

Name Published In: Minnesota Bot. Stud. 1: 815 (1897)

Other References

Wagner et al. 1990:1309 (K)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN David H. Lorence 5948 Kauai PTBG 4/10/1988
2 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Metrosideros polymorpha Elfin Forest. PRESERVED_SPECIMEN Jesse Adams JWA_28 Kauai PTBG 10/13/2013
3 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Metrosideros open forest 2-10% cover @ 7-10 m tall. PRESERVED_SPECIMEN K. R. Wood 12033 Kauai PTBG 8/10/2006
4 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN K. R. Wood 184 Kauai PTBG 9/10/1988
5 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN K. R. Wood 3544 Kauai PTBG 9/19/1994
6 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN K. R. Wood 3585 Kauai PTBG 9/26/1994
7 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN K. R. Wood 7326 Kauai PTBG 5/27/1998
8 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Closed Metrosideros forest, 8-12 m canopy. PRESERVED_SPECIMEN K. R. Wood 13258 Kauai PTBG 9/10/2008
9 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Metrosideros-Dicranopteris lowland wet shrubland with small pockets of mixed wet forest below ridgeline. PRESERVED_SPECIMEN K. R. Wood 15234 Kauai PTBG 9/21/2012
10 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN K. R. Wood 2853 Kauai PTBG 11/10/1993
11 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN K. R. Wood 3600 Kauai PTBG 9/26/1994
12 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Wagner, W. L.; Takeuchi, W. N.; Imada, C. T. 6051 Kauai US 4/10/1988
13 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN w. l. wagner, d. h. lorence, r. hanford & k. wood 6292 Kauai US 11/21/1989
14 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information On bank of intermittent stream PRESERVED_SPECIMEN Bryan, E. H. 1459 Kauai US 7/29/1940
15 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Wood, K. R.; Perlman, S. P. 3585 Kauai US 9/26/1994
16 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Streamside PRESERVED_SPECIMEN Tangalin, N.; Trauernicht, C.; Emmonds, M.; Griffin-Noyes, E. 1812 Kauai US 10/1/2008
17 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Wagner, W. L.; Lorence, D. H.; Hanford, R.; Wood, K. 6292 Kauai US 11/21/1989
18 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN a. a. heller 2852 Kauai US 1895-09-10
19 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN a. a. heller 2852 Kauai US 1895-10-01
20 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN n. tangalin, c. trauernicht, m. emmonds & e. griffin-noyes 1812 Kauai US 10/1/2008
21 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Heller, A. A. 2852 Kauai US 1895-00-00
22 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN t. w. flynn 3392 Kauai US 5/11/1989
23 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN a. s. hitchcock 15365 Kauai US 10/20/1916
24 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN k. r. wood & s. p. perlman 3585 Hawaii US 9/26/1994
25 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN w. l. wagner, w. n. takeuchi & c. imada 6051 Kauai US 4/10/1988
26 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Flynn, T. W. 3392 Kauai US 5/11/1989
27 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information On bank of intermittent stream PRESERVED_SPECIMEN e. bryan 1459 Kauai US 7/29/1940
28 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Metrosideros-Dicranopteris lowland wet shrubland with small pockets of mixed wet forest below ridgeline with Broussaisa arguta, Melicope feddei, Psychotria mariniana & P. wawrae, Kadua affinis, Antidesma platyphylla, Alyxia stellata, Syzygium sandwicensis, Dubautia laxa PRESERVED_SPECIMEN wood, k.r. query, m. collector number: 15234.00 Kauai BISH 9/21/2012
29 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Forbes, C. N. 86.K. Kauai US 7/23/2009
30 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Forbes, C.N. 86K Kauai BISH 7/23/1909
31 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN christensen, c. collector number: 86 Kauai BISH 10/23/1976
32 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Woods. PRESERVED_SPECIMEN st.john, h. collector number: 10692 Kauai BISH 12/26/1930
33 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Steve Perlman 18741 Kauai PTBG 1/7/2004
34 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Montane Metrosideros Riparian zone PRESERVED_SPECIMEN Natalia Tangalin 2498 Kauai PTBG 2/3/2011
35 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN herbst, d.r. collector number: 2224 Kauai BISH 11/3/1971
36 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN rock, j.f.c. collector number: 1694 Kauai BISH 2/14/1909
37 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN forbes, c.n. collector number: 101h Hawaii BISH 7/31/1909
38 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Rock, J.F.C. 5214 Kauai BISH 8/1/1909
39 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Rock, J.F.C. 7148 Kauai BISH
40 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Along stream bank. PRESERVED_SPECIMEN flynn, t. collector number: 3392 Kauai BISH 5/11/1989
41 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Chock, A.K. 995 Kauai BISH 8/19/1953
42 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Wet forest along stream. PRESERVED_SPECIMEN St. John, H. 13795 Kauai BISH 12/28/1933
43 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Forbes, C.N. 101h Kauai BISH 7/31/1909
44 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Christensen, C. 86 Kauai BISH 10/23/1976
45 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron montane wet forest, bogs and riparian habitats w/ Broussaisia arguta, Dubautia paleata, Labordia waialealae, L. Pumila, Polyscias waialealae, Vaccinum calycinum, Pipturus ruber, Deschampsia nubigena, Machaerina angustifolia, Melicope clusiifolia, PRESERVED_SPECIMEN Wood, K.R. 17906 Kauai BISH 8/13/2018
46 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information upper drainage, closed to open mixed mesic forest, talus slopes; Metrosideros, Diospyros, Cheirodendron fauriei, Antidesma platyphylla, Pouteria sandwicensis, Pleomele aurea, Dodonaea viscosa, Pritchardia minor, Microlepia strigosa, Diplazium sandwichianum, Dryopteris fusco-atra PRESERVED_SPECIMEN Wood, K.R. 12141 Kauai BISH 9/17/2006
47 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN forbes, c.n. collector number: 563k Oahu BISH 10/10/1916
48 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron-Dicranopteris wet forest and deeply carved drainage with riparian vegetation Kadua affinis, Psychotria mariniana, P. hexandra, Coprosma kauaiensis, Syzygium sandwicensis, Perrottetia sandwicensis, Dubautia raillardioides, D. laxa, Melicope wawrana, M. clusiifolia, Myrsine wawr PRESERVED_SPECIMEN Wood, K.R. Query, M. Collector Number: 15269.00 Kauai BISH 11/7/2012
49 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN forbes, c.n. collector number: 115k Kauai BISH 8/2/1909
50 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Open healthy forest; primary vegetation: Metrosideros polymorpha (sometimes pubescent), Ilex anomala, Myrsine lessertiana, Cheirodendron trigynum, Coprosma ochracea & Cibotium glaucum. Soil a shallow well drained organic found over an a`a flow. PRESERVED_SPECIMEN wood, k.r. boynton, d. collector number: 11524 Hawaii BISH 9/9/2005
51 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Rainforest. PRESERVED_SPECIMEN MacDaniels, L.H. 860 Kauai BISH 2/22/1927
52 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Lowland Dry Forest at 1350 ft on a north facing slope with Erythrina, Diospyros, Canthium, Leucaena, Achyranthes, Sida, Alyxia, Plumbago, Chenopodium, Myoporum, Peperomia, and an assortment of alien grasses and herbs PRESERVED_SPECIMEN stone, b.c. collector number: 1490 Maui BISH 12/26/1956
53 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Metrosideros-Dicranopteris lowland wet shrubland with small pockets of mixed wet forest below ridgeline with Broussaisa arguta, Melicope feddei, Psychotria mariniana & P. wawrae, Kadua affinis, Antidesma platyphylla, Alyxia stellata, Syzygium sandwicensis, Dubautia laxa PRESERVED_SPECIMEN Wood, K.R. 15234 Kauai BISH 9/21/2012
54 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN wood, k.r. kirkpatrick, m., perlman, s. collector number: 15551 Oahu BISH
55 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information on branch of Metrosideros PRESERVED_SPECIMEN christensen, c. collector number: 260 Oahu BISH 7/16/1977
56 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Montane Metrosideros Riparian zone W/Melicope clusifolia, Broussaisia, Pepperomia, Asplenium, Mosses, Selaginella, Metrosideros, Rubus rosifolius, Stenogyne purpurea, Sadleria, Cibotium, Labordia waialealae, Dubautia, Pipturus, Lobel(label end) PRESERVED_SPECIMEN Tangalin, N. 2498 Kauai BISH 2/3/2011
57 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN heller, a.a. collector number: 2852 Kauai BISH 1895-09-02
58 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN rock, j.f.c. collector number: 1672 Kauai BISH 2/14/1909
59 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Closed Metrosideros forest, 8-12 m canopy. Understory dominated by Antidesma w/ mixed wet associates, Syzygium, Broussaisia, Perrottetia, Cyrtandra spp, Psychotria spp, Dubautia spp, Labordia spp, Coprosma waimeae, Cheirodendron spp, Tetraplasandra PRESERVED_SPECIMEN Wood, K.R. 13258 Kauai BISH 9/10/2008
60 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN rock, j.f.c. collector number: 1725 Kauai BISH
61 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Common along stream. PRESERVED_SPECIMEN Wagner, W.L. 6292 Kauai BISH 11/21/1989
62 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Wet forest along stream. PRESERVED_SPECIMEN st.john, h. fosberg, f.r. collector number: 13795 Kauai BISH 12/28/1933
63 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN rock, j.f.c. collector number: 7148 Kauai BISH
64 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information growth, except upper part of valley dominated by Metrosideros with understory of Psychotria, Hedyotis terminalis, Cyanea (3 species), Dicranopteris (in open areas), 3 spp. of Cyrtandra, Labordia hirtella (in upper part). Dense, wet forest, along stream. Moderate to heavy Psidium PRESERVED_SPECIMEN Wagner, W.L. 6051 Kauai BISH 4/10/1988
65 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information in savannah with Sapindus. About 1.8 m. from ground on slightly flaking bark of Sapindus. Common. PRESERVED_SPECIMEN forbes, c.n. collector number: 101g Hawaii BISH 7/31/1909
66 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN rock, j.f.c. collector number: 2370 Kauai BISH 2/14/1909
67 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Krajina, V.J. s.n. Kauai BISH 2/6/1930
68 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Common in this area; also seen along forestry roads above the experimental station. PRESERVED_SPECIMEN wagner, w.l. lorence, d.; hanford, r.; wood, k. collector number: 6292 Hawaii BISH 11/21/1989
69 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Wet forest. PRESERVED_SPECIMEN St. John, H. 13701 Kauai BISH 12/27/1933
70 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Montane wet forest with 60% cover of Metrosideros at 9 m tall and 30% cover of Cheirodendron at 7 m tall, diverse understory with Melicope clusiifolia, M. wawrana, Vaccinium calycinum, Broussaisia arguta, Trematolobelia kauaiensis, Lobelia hypoleuca, rich fern understory with Athyrium microphyllum, Sadleria pallida, Asplenium polyodon, abundant terrestrial and epiphytic moss. PRESERVED_SPECIMEN Wood, K.R. 11524 Kauai BISH 9/9/2005
71 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN c. n. forbes 86.k. Kauai US 7/23/1909
72 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN forbes, c.n. collector number: 101f Kauai BISH 7/31/1909
73 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Heller, A. A. 2852 Kauai US 1895-00-00
74 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information On bank of intermittent stream. PRESERVED_SPECIMEN Bryan Jr., E.H. 1459 Kauai BISH 7/29/1940
75 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Forbes, C.N. 101g Kauai BISH 7/31/1909
76 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information mixed forest with riparian vegetation Pisonia umbellifera, Antidesma platyphylla Syzygium sandwicensis, Psychotria mariniana, P. mauiensis, P. wawrae, Melicope clusiifolia, Broussaisia arguta, Cyrtandra kealiae urceolata, C. longifolia, C PRESERVED_SPECIMEN Wood, K.R. 15551 Kauai BISH 6/27/2013
77 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Along stream bank. PRESERVED_SPECIMEN Flynn, T. 3392 Kauai BISH 5/11/1989
78 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Along stream bank in low, open wet forest with Metrosideros polymorpha (dominant), Antidesma, Syzygium, invaded by Psidium cattleianum. PRESERVED_SPECIMEN lorence, d.h. flynn, t.; wood, k.r; ragone, d. collector number: 6653 Kauai BISH 12/18/1990
79 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron-Dicranopteris wet forest and deeply carved drainage with riparian vegetation Kadua affinis, Psychotria mariniana, P. hexandra, Coprosma kauaiensis, Syzygium sandwicensis, Perrottetia sandwicensis, Dubautia raillardioides, D. laxa, Melicope wawrana, M. clusiifolia, Myrsine wawr PRESERVED_SPECIMEN Wood, K.R. 15269 Kauai BISH 11/7/2012
80 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Along stream bank in low, open wet forest with Metrosideros polymorpha (dominant), Antidesma, Syzygium, invaded by Psidium cattleianum. PRESERVED_SPECIMEN Lorence, D.H. 6653 Kauai BISH 12/18/1990
81 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information On bank of intermittent stream. PRESERVED_SPECIMEN bryan jr., e.h. collector number: 1459 Kauai BISH 7/29/1940
82 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN krajina, v.j. collector number: s.n. Kauai BISH 2/6/1930
83 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Heller, A.A. 2852 Kauai BISH 1895-00-00
84 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information In open sun. PRESERVED_SPECIMEN Christensen, C. 260 Kauai BISH 7/16/1977
85 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN rock, j.f.c. collector number: 5214 Kauai BISH 8/8/1909
86 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Rock, J.F.C. 1725 Kauai BISH
87 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information In wet forest. PRESERVED_SPECIMEN Stone, B.C. 1490 Kauai BISH 12/26/1956
88 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information calm waters PRESERVED_SPECIMEN heller, a.a. collector number: 2852 Maui BISH 1895-09-02
89 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Forbes, C.N. 115K Kauai BISH 8/2/1909
90 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information In a more protected drainage, on the mid slope of a shallow gulch. Vegetation: Cheirodendron platyphyllum, Cheirodendron trigynum, Metrosideros polymorpha, Metrosideros rugosa, Broussaisia arguta, Hedyotis fosbergii, Dicranopteris linearis, Machaerina angustifolia. PRESERVED_SPECIMEN chock, a.k. collector number: 995 Oahu BISH 8/19/1953
91 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN wagner, w.l. takeuchi, w.; imada, c. collector number: 6051 Hawaii BISH 4/10/1988
92 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Rock, J.F.C. 2370 Kauai BISH 2/14/1909
93 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN forbes, c.n. collector number: 86k Kauai BISH 7/23/1909
94 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Rock, J.F.C. 1615 Kauai BISH 3/3/1909
95 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Closed Metrosideros forest, 8-12 m canopy. Understory dominated by Antidesma w/ mixed wet associates, Syzygium, Broussaisia, Perrottetia, Cyrtandra spp, Psychotria spp, Dubautia spp, Labordia spp, Coprosma waimeae, Cheirodendron spp, Tetraplasandra PRESERVED_SPECIMEN wood, k.r. collector number: 13258 Kauai BISH 9/10/2008
96 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Rock, J.F.C. 1694 Kauai BISH 2/14/1909
97 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Woods. PRESERVED_SPECIMEN St. John, H. 10692 Kauai BISH 12/26/1930
98 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN rock, j.f.c. collector number: 1615 Kauai BISH 3/3/1909
99 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Wet forest. PRESERVED_SPECIMEN st.john, h. fosberg, f.r. collector number: 13701 Kauai BISH 12/27/1933
100 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN a. a. heller 2852 A
101 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information On slopes of losse rock and rubble. A 6 m tree with canopy associates Streblus, Elaeocarpus, and Planchonella. Understory elements are Embelia, Neraudia, Urera, and Claoxylon. PRESERVED_SPECIMEN heller, a.a. collector number: 2852 Oahu BISH 1895-09-02
102 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Herbst, D.R. 2224 Kauai BISH 11/3/1971
103 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Forbes, C.N. 101f Kauai BISH 7/31/1909
104 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron-Dicranopteris wet forest and deeply carved drainage with riparian vegetation Kadua affinis, Psychotria mariniana, P. hexandra, Coprosma kauaiensis, Syzygium sandwicensis, Perrottetia sandwicensis, Dubautia raillardioides, D. laxa, Melicope wawrana, M. clusiifolia, Myrsine wawr PRESERVED_SPECIMEN wood, k.r. query, m. collector number: 15269.00 Kauai BISH 11/7/2012
105 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Rainforest. PRESERVED_SPECIMEN macdaniels, l.h. collector number: 860 Kauai BISH 2/22/1927
106 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information ad m. quovours [?] PRESERVED_SPECIMEN a. a. heller 2852 FH
107 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Heller, A. A. (Coll. Delessert) G 1895-09-24
108 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Heller, A. A. (Coll. Barbey-Boissier) G 1895-09-24
109 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Small tree at edge of ravine, in full flower (red and yellow, pink bracts). PRESERVED_SPECIMEN heller, a.a. 2852 LSU 1895-10-01
110 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN amos arthur heller; b.1867; d.1944; a.heller 2852 NHMUK 1895-10-01
111 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN k. r. wood 15269 NY 11/7/2012
112 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN K. R. Wood 15269 NY 11/7/2012
113 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Tim Flynn 6191 Kauai PTBG 6/26/1997
114 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN K. R. Wood 1588 Kauai PTBG 1/23/1992
115 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Dicranopteris-Metrosideros Windward Wet Forest PRESERVED_SPECIMEN Natalia Tangalin 2528 Kauai PTBG 2/24/2011
116 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Metrosideros mixed wet forest with riparian vegetation. PRESERVED_SPECIMEN K. R. Wood 15551 Kauai PTBG 6/27/2013
117 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Steve Perlman 17272 Kauai PTBG 8/10/2000
118 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Tim Flynn 3392 Kauai PTBG 5/11/1989
119 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN K. R. Wood 3533 Kauai PTBG 9/19/1994
120 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN K. R. Wood 11524 Kauai PTBG 9/9/2005
121 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Low, open wet forest PRESERVED_SPECIMEN David H. Lorence 6653 Kauai PTBG 12/18/1990
122 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information upper drainage, closed to open mixed mesic forest, talus slopes; Metrosideros, Diospyros, Cheirodendron fauriei, Antidesma platyphylla, Pouteria sandwicensis, Pleomele aurea, Dodonaea viscosa, Pritchardia minor, Microlepia strigosa, Diplazium sandwichianum, Dryopteris fusco-atra PRESERVED_SPECIMEN wood, k.r. tangalin, n. collector number: 12141 Kauai BISH 9/17/2006
123 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Derral Herbst 2224 Kauai PTBG 11/3/1971
124 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron-Dicranopteris wet forest and deeply carved drainage with riparian vegetation. PRESERVED_SPECIMEN K. R. Wood 15269 Kauai PTBG 11/7/2012
125 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information almost undisturbed low elevation rain forest. PRESERVED_SPECIMEN Tim Flynn 2656 Kauai PTBG 12/10/1987
126 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Uma Nagendra UJN002 Kauai PTBG 10/24/2017
127 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Gulch on north-east facing slopes PRESERVED_SPECIMEN Seana Walsh SKW674 Kauai PTBG 2/5/2019
128 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Clay Trauernicht 571 Kauai PTBG 10/1/2008
129 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN K. R. Wood 12141 Kauai PTBG 9/17/2006
130 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN K. R. Wood 5280 Kauai PTBG 6/5/1996
131 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Metrosideros-Cheirodendron montane wet forest, bogs and riparian habitats PRESERVED_SPECIMEN K. R. Wood 17906 Kauai PTBG 8/13/2018
132 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Forbes, C.N. 563K Kauai BISH 10/10/1916
133 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Rock, J.F.C. 1672 Kauai BISH 2/14/1909
134 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information Growing on trail in mesic forest PRESERVED_SPECIMEN a. a. heller 2852 NY 1895-10-01
135 Pipturus ruber A.Heller Contact Bishop Museum Botany department for locality information PRESERVED_SPECIMEN Hitchcock, A. S. 15365 Kauai US 10/20/2016
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